Fossils of a whale that beached on an African shore more than a million years ago and was subsequently butchered by hominids have been recovered by paleontologists.

A. that beached on an African shore more than a million years ago and was subsequently butchered by hominids have

B. that beached on an African shore more than a million years ago and then was subsequently butchered by hominids has

C. that beached on an African shore more than a million years ago, which was subsequently butchered by hominids, has

D. having been beached on an African shore more than a million years ago and subsequently butchered by hominids, have

E. having beached on an African shore more than a million years ago and then subsequently were butchered by hominids have